Chapter 17

•      Plant Form and Function

With emphasis on Angiosperms, the flowering plants

 

Plants are

•      Multicellular eukaryotes that have cellulose rich cell walls and use starch as an energy reserve.

•      Most are photosynthetic autotrophs

•      Have alternation of generations

–   Sporophyte

–   gametophyte

 

Plants are classified by

•      Presence or absence of vessels and seeds

•      Lack vessels (and seeds): Bryophytes

•      Have vessels: Vascular plants

•      Vascular plants

–   Without seeds: Ferns and fern allies

–   With seeds:

•     gymnosperms (naked seeded)

•    angiosperms (covered seeds, the flowering plants)

Plant Evolution

•     Plants have a long evolutionary history

Evolutionary trends

•      Characteristics shared with green algae:

·   Chlorophyll a is the primary photosynthetic pigment, chlorophyll b and carotenoids are accessory pigments.

·   Starch is the reserve food, deposited in chloroplasts.

·   Cellulose cell walls in all plants (& some green algae).

·   In cell division, form phragoplast and cell plate in all plants (& some green algae)

•       This evidence suggests that green plants arose from some green algae.

Plants are important for

•      Food

•      Wood -fuel, building, furniture, etc

•      Fiber - clothing, paper, etc.

•      Drugs (25% of prescription drugs)

•      Oxygen production

Success of the Angiosperms

•      The angiosperms are seed-bearing vascular plants

•      In terms of distribution and diversity, they are the most successful plants on Earth

•      The structure and function of this plant group help explain its success

 

Flowering Plants Can Be divided into two Groups: Monocots and Dicots

How Are Plant Bodies Organized?

•       Flowering Plants Have a Root System and a Shoot System

Shoots and Roots

•      Shoots

–   Produce food by photosynthesis

–   Carry out reproductive functions

–   Transport materials

•      Roots

–   Anchor the plant

–   Penetrate the soil and absorb water and dissolved minerals

–   Store food

Different Processes Are Responsible for Growth in Length and Width

•      Meristems are region where cell divisions produce plant growth

•      Apical meristems

–   Lengthen stems and roots

–   Responsible for primary growth

•      Lateral meristems

–   Increase width of stems

–   Responsible for secondary growth

Apical Meristems

•      Lengthen shoots and roots from tips

•      During growth, meristem cells  give rise to differentiated cells

 

Lateral Meristems

•      Increases girth of older roots and stems

•      Cylindrical arrays of cells

Three Tissue Types

 

What Are the Tissues  and Cell Types of Plants?

•      Dermal Tissue Covers the Plant Body

Epidermis:

•      Covers and protects plant surfaces

•      Secretes a waxy, waterproof cuticle

 

What Are the Tissues  and Cell Types of Plants?

•       Ground Tissue Makes Up Most of the Young Plant Body

–    Carry out most function

•    Photosynthesis

•    Store sugars, starches

•    Secrete hormones

•    Provide support

 

 

What Are the Tissues  and Cell Types of Plants?

•      Vascular Tissue Consists of Xylem and Phloem

–   Xylem Conducts Water and Dissolved Nutrients from the Roots to the Rest of the Plant

–   Phloem Conducts Substances throughout the Plant

 

 Xylem

 

Phloem:
A Complex Vascular Tissue

•      Transports sugars 

•      Main conducting cells are sieve-tube members

•      Companion cells assist in the loading of sugars

 

 

 

Root Structure

•       Root cap covers tip

•       Apical meristem produces the cap

•       Cell divisions at the apical meristem cause the root to lengthen

•       Farther up, cells differentiate and mature

Internal Structure of a Root

•       Outermost layer is epidermis

•       Root cortex is beneath the epidermis

•       Endodermis surrounds the vascular cylinder

•       In some plants, there is a central pith

How Do Roots Grow and What Do They Do?

•      The Epidermis of the Root Is Very Permeable to Water

 

How Do Roots Grow and What Do They Do?

•      The Cortex Controls the Absorption of Water and Nutrients

–   Outer layer is loose cells

–   Inner layer is endodermis

Function of Endodermis

•      Ring of cells surrounds vascular cylinder

•      Cell walls are waterproof 

•      Water can only enter vascular cylinder by moving through endodermal cells

•      Allows plant to control inward flow

 

Internal Structure of a
Dicot Stem

•      Outermost layer is epidermis

•      Cortex lies beneath epidermis

•      Ring of vascular bundles separates the cortex from the pith

•      The pith lies in the center of the stem

Internal Structure of a Monocot Stem

How Do Stems Grow and What Do They Do?

•      The Epidermis of the Stem Retards Water Loss While Allowing Carbon Dioxide to Enter

•      The Cortex and Pith Support the Stem, Store Food, and Photosynthesize

•      Vascular Tissues in Stems Transport Water, Dissolved Nutrients, and Hormones

 

 

Herbaceous vs. Woody

•      Herbaceous have primary growth, live a single year – grass, beans

•      Woody- have secondary growth, can live multiple years – trees and shrubs

 

•      Secondary Growth Produces Thicker, Stronger Stems

 

Secondary Growth

•      Occurs in all gymnosperms, some monocots, and many dicots

•      A ring of vascular cambium produces secondary xylem and phloem

•      Wood is the accumulation of these secondary tissues, especially xylem

 

 

Secondary Growth

•       Lateral meristem

•       One or two cells thick

•       Divides in two directions

•       Cells formed to the outside become secondary phloem

•       Cells formed to the inside become secondary xylem

Woody Stem

Annual Rings

•      Concentric rings of secondary xylem

•      Alternating bands of early and late wood

•      Early wood

–   Xylem cells with large diameter, thin walls

•      Late wood

–   Xylem cells with smaller diameter, thicker walls

How Do Stems Grow and What Do They Do?

Secondary Growth Causes the Epidermis to Be Replaced by Woody Cork

 

 

 

Some Specialized Stems Produce New Plants or Store Water or Food

What Is the Structure of Leaves and What Do They Do?

•      Leaves Have Two Major Parts:

–   Stalk

–   Blade

•      Photosynthesis, gas exchange, transport

 

Gross Leaf Structure

Adapted for Photosynthesis

•      Leaves are usually thin

–   High surface area-to-volume ratio

–   Promotes diffusion of carbon dioxide in, oxygen out

•      Leaves are arranged to capture sunlight

–   Are held perpendicular to rays of sun

–   Arrange so they don’t shade one another

 

Leaf Veins: Vascular Bundles

•       Xylem and phloem; often strengthened with fibers

•       In dicots, veins are netlike

•       In monocots, they are parallel

 

•       Specialized Leaves May Provide Support, Store Food, or Even Capture Insects

 

How Do Plants Acquire  Nutrients?

•      Roots Take Up Minerals Dissolved in Water

•      Fungi and Bacteria Help Plants Acquire Nutrients

–   Fungal Mycorrhizae Help Plants Acquire Minerals

–   Bacteria-Filled Nodules on Their Roots Help Legumes Acquire Nitrogen

 

 

How Do Plants Acquire Water and Transport Water and Minerals?

•      Water Enters Roots Mainly by Pressure Differences Created by Transpiration

•      Adjustable Stomata Control the Rate of Transpiration

 

•      Transpiration Provides the Force for Water Movement in Xylem

 

 

Plant diversity

•      Even plants that are “weeds” may contain drugs or genes that might be useful …

•      We need to preserve habitats as well as fund seed banks and other conservation efforts.

 

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